1. School of Biological Sciences; University of Canterbury; PB 4800 Christchurch New Zealand
2. Nigerian Montane Forest Project; Yelwa village Taraba State Nigeria
3. Department of Biology; Gombe State University; Tudan Wada Street Gombe Gombe State Nigeria
4. Department of Math and Statistics; University of Canterbury; PB 4800 Christchurch New Zealand
5. Département Ecologie et Gestion de la Biodiversité; Muséum National d'Histoire Naturelle; CNRS UMR 7179; 1 avenue du Petit Château 91800 Brunoy France