1. Department of Zoology; University of Oxford; Tinbergen Building, South Parks Road Oxford OX1 3PS UK
2. School of Geography and the Environment; University of Oxford; South Parks Road Oxford OX1 3QY UK
3. United Nations Environment Programme World Conservation Monitoring Centre; 219 Huntingdon Road Cambridge CB3 0DL UK
4. Department of Geography; University of Cambridge; Downing Place Cambridge CB2 3EN UK
5. Department of Genetics, Evolution and Environment, Centre for Biodiversity and Environment Research; University College London; Gower Street London WC1E 6BT UK