1. Division of Forest and Biomaterial Science Graduate School of Agriculture Kyoto University Kitashirakawa OiwakeKyoto Japan
2. Department of Plant Sciences Faculty of Agriculture Rajarata University Anuradhapura Sri Lanka
3. Plant Ecophysiology & Evolution Group Guangxi Key Laboratory for Forest Ecology and Conservation College of Forestry Guangxi University NanningGuangxi China
4. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐Bioresources Guangxi University Nanning Guangxi China
5. Smithsonian Tropical Research Institute Balboa Panama