1. Department of Environment, Earth and Ecosystems; Open University; Walton Hall; Milton Keynes; MK7 6AA; UK
2. Department of Mathematics and Statistics; Open University; Walton Hall; Milton Keynes; MK7 6AA; UK
3. Institute of Systematic Botany; University of Zurich; Xxxxx; Switzerland
4. Kirstenbosch Research Centre; South African National Biodiversity Institute; Xxxxxxx; South Africa