1. Division of Biological Sciences Section Cell & Developmental Biology University of California San Diego 9500 Gilman Drive La Jolla CA 92093 USA
2. Laboratoire de Recherche en Sciences Végétales CNRS, UPS Université de Toulouse 24 chemin de Borde Rouge, Auzeville, BP42617 Castanet Tolosan 31326 France
3. Department of Plant and Soil Sciences University of Kentucky Lexington KY 40546‐0091 USA