1. Department of Ecology; Evolution and Natural Resources and Institute of Marine and Coastal Science; Rutgers University; New Brunswick New Jersey 08901 USA
2. Unité Mixte de Recherche; Centre National de la Recherche Scientifique 7138, Systématique, Adaptation, Evolution; Université Pierre et Marie Curie; Paris 75005 France
3. Department of Ecology; Evolution Natural Resources; Rutgers University; New Brunswick New Jersey 08901 USA
4. Department of Biological Sciences; Sungkyunkwan University; Suwon 440-746 Korea