1. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture; Wuhan Botanical Garden; Chinese Academy of Sciences; Wuhan Hubei China
2. University of Chinese Academy of Sciences; Beijing China
3. Department of Biology; Oberlin College; Oberlin Ohio USA
4. The State Key Laboratory of Species Identification; Yunnan Entry-Exit Inspection and Quarantine Bureau; Kunming Yunnan China
5. Key Laboratory of Biodiversity and Biogeography; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan China