1. Institute of Biological Environmental and Rural Sciences (IBERS) Aberystwyth, University Aberystwyth SY23 3DA UK
2. Cell and Molecular Sciences The James Hutton Institute (JHI) Invergowrie Dundee DD2 5DA UK
3. Information and Computational Sciences The James Hutton Institute (JHI) Invergowrie Dundee DD2 5DA UK
4. Division of Plant Sciences University of Dundee at The James Hutton Institute Invergowrie Dundee DD2 5DA UK