1. Division of Plant Sciences, College of Life Sciences, University of Dundee at The James Hutton Institute, Invergowrie, Dundee DD2 5DA, United Kingdom
2. School of Biosciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, United Kingdom
3. The James Hutton Institute, Invergowrie, Dundee DD2 5DA, United Kingdom