1. Laboratory of Animal Breeding and Genetics; Graduate School of Agricultural Science; Kobe University; Kobe Hyogo Japan
2. National Livestock Breeding Center; Nishigo Fukushima Japan
3. National Institute of Livestock and Grassland Science; Tsukuba Ibaraki Japan
4. Gifu Prefectural Livestock Research Institute; Takayama Gifu Japan
5. Food Resources Education & Research Center; Kobe University; Kasai Hyogo Japan