1. Department of Biology, FFCLRP; University of Sao Paulo, Ribeirão Preto; São Paulo Brazil
2. Ecology and Evolution, Department of Biological Sciences; University of Illinois; Chicago IL USA
3. Department of Soils and Fertilizers, FCAV; São Paulo State University; São Paulo Brazil
4. Federal Institute Goiano, Rio Verde Campus; Goiás Brazil
5. Institute of Botany; São Paulo Brazil