1. Plymouth Marine Laboratory Prospect Place Plymouth PL1 3DH UK
2. Institute of Marine Sciences University of California Santa Cruz CA 95064 USA
3. Marine Biology and Ecology Research Centre Plymouth University Plymouth PL4 8AA UK
4. The Laboratory Marine Biological Association of the United Kingdom Citadel Hill Plymouth PL1 2PB UK
5. Ocean and Earth Science National Oceanography Centre Southampton Waterfront Campus University of Southampton Southampton SO14 3ZH UK
6. Centre for Biological Sciences Building 85 Highfield Campus University of Southampton Southampton SO17 1BJ UK