1. Department of Zoology and Entomology Rhodes University Grahamstown South Africa
2. CCMAR, CIMAR Associated Laboratory University of Algarve Faro Portugal
3. Laboratoire d'Océanologie et de Géosciences Univ. LilleCNRSUniv. Littoral Côte d'OpaleUMR 8187 LOG Lille France
4. Department of Marine Resources and Energy Tokyo University of Marine Science and Technology Tokyo Japan