1. Department of Ecology and Evolutionary Biology University of Arizona 1041 East Lowell, Tucson AZ 85721 USA
2. Departamento de Biología Recinto Universitario de Mayagüez Universidad de Puerto Rico Mayagüez Puerto Rico 00681 USA
3. CEFE UMR 5175 CNRS Université de Montpellier Université Paul‐Valéry Montpellier EPHE‐1919 route de Mende F‐34293 Montpellier, Cedex 5 France
4. The Santa Fe Institute University of Arizona 1399 Hyde Park Road Santa FeNM 87501 USA