1. Department of ZoologyUniversity of Oxford Oxford UK
2. Evolutionary Biodemography LaboratoryMax Planck Institute for Demographic Research Rostock Germany
3. Centre for Biodiversity and Conservation ScienceUniversity of Queensland St Lucia Qld Australia
4. CEFE, CNRSUniv MontpellierUniv Paul Valéry Montpellier 3, EPHE, IRD Montpellier France
5. Department of Animal & Plant SciencesThe University of Sheffield Sheffield UK