1. Department of Wildland Resources and the Ecology Center; Utah State University; Logan UT 84322 USA
2. Centro de Investigación en Ecosistemas de la Patagonia (CIEP) Conicyt-Regional R10C1003; Universidad Austral de Chile; Ignacio Serrano 509 Coyhaique Chile
3. Department of Biology; University of Maryland; College Park MD 20742 USA