1. Key Laboratory of Agro‐ecological Processes in Subtropical Region Institute of Subtropical Agriculture Chinese Academy of Sciences Changsha China
2. State Key Laboratory of Biocontrol School of Ecology Sun Yat‐sen University Guangzhou China
3. Huanjiang Observation and Research Station for Karst Ecosystems Chinese Academy of Sciences Huanjiang China
4. South China Botanical Garden Chinese Academy of Sciences Guangzhou China
5. Key Laboratory of Environment Change and Resources Use in Beibu Gulf Ministry of Education Nanning Normal University Nanning China