1. Laboratoire de Biométrie et Biologie Evolutive UMR 5558, CNRS, Université Lyon 1 Université de Lyon Villeurbanne France
2. ESE, Ecology and Ecosystem Health, Agrocampus Ouest INRA Rennes France
3. LBLGC EA 1207 INRA, Université d'Orléans, USC 1328 Orléans France
4. Department of Biology and Graduate Degree Program in Ecology Colorado State University Fort Collins Colorado