1. Evolution & Ecology Research Centre; University of New South Wales; Sydney; NSW 2052; Australia
2. Department of Biology; University of Antwerp; Universiteitsplein 1; B-2610; Wilrijk; Belgium
3. School of Biological Sciences; The University of Sydney; NSW 2010; St Lucia; Australia
4. School of Biological Sciences; The University of Queensland; QLD 4072; St Lucia; Australia