1. School of Marine Science and Engineering Qingdao Agricultural University Qingdao China
2. State Key Laboratory of Freshwater Ecology and Biotechnology Institute of Hydrobiology Chinese Academy of Sciences Wuhan China
3. Key Laboratory of Maricultural Organism Disease Control Ministry of Agriculture and Rural Affairs Yellow Sea Fisheries Research Institute Chinese Academy of Fishery Sciences Qingdao China
4. Laboratory for Marine Biology and Biotechnology Pilot National Laboratory for Marine Science and Technology Qingdao China