1. Department of Biological Sciences; Graduate School of Science; The University of Tokyo; 7-3-1 Hongo, Bunkyo-ku Tokyo 113-0033 Japan
2. Graduate School of Biostudies; Kyoto University; Yoshida Konoecho; Sakyo-ku Kyoto 606-8501 Japan
3. Plant Global Education Project; Graduate School of Biological Sciences; Nara Institute of Science and Technology; 8916-5 Takayama Ikoma Nara 630-0192 Japan
4. Departamento de Genética Molecular de Plantas; Centro Nacional de Biotecnología; Consejo Superior de Investigaciones Científicas; Madrid 28049 Spain