1. Department of Ecology, Evolution, and Behavior University of Minnesota Twin Cities Saint Paul MN 55108 USA
2. Department of Fisheries, Wildlife, and Conservation Biology University of Minnesota Twin Cities Saint Paul MN 55108 USA
3. Centre for Biodiversity Theory and Modelling, Theoretical and Experimental Ecology Station CNRS and Paul Sabatier University 09200 Moulis France
4. Department of Forest Resources University of Minnesota Twin Cities Saint Paul MN 55108 USA
5. Hawkesbury Institute for the Environment Western Sydney University Penrith NSW 2753 Australia
6. Department of Biology McGill University Montreal QC H3A 1B1 Canada
7. Department of Plant Sciences University of Oxford Oxford OX1 3RB UK
8. URPP Global Change and Biodiversity Department of Evolutionary Biology and Environmental Studies University of Zürich Winterthurerstrasse 190 8057 Zürich Switzerland