1. Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, UK
2. College of Resources and Environmental Science, Hebei Agricultural University, Baoding, Hebei Province 071001, China
3. Institute for Environmental Genomics, University of Oklahoma, 101 David L. Boren Blvd, Norman, OK 73019, USA
4. Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 3UU, UK
5. Department of Disease and Stress Biology, John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK