1. The Sainsbury Laboratory, John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK
2. Department of Plant Pathology, Physiology and Weed Science, Virginia Tech, Latham Hall, Blacksburg VA 24061, USA
3. Centre for the Analysis of Genome Evolution and Function, University of Toronto, 25 Willcocks Street, Toronto, Ontario M5S 3B2, Canada