1. Department of Biological Sciences, Macquarie University, North Ryde, NSW 2109, Australia
2. Grantham Institute and Division of Biology, Imperial College, Silwood Park, Ascot SL5 7PY, UK
3. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Xiangshan Nanxincun 20, 100093 Beijing, China
4. School of Geographical Sciences, University of Bristol, Bristol BS8 1SS, UK
5. Alfred Wegener Institute for Polar and Marine Research, Telegrafenberg A43, D‐14473 Potsdam, Germany