1. Graduate School of Life Sciences, Tohoku University, 6‐3 Aoba, Sendai 980‐8578, Japan;
2. Present address: Center for Bioresource Field Science, Kyoto Institute of Technology, Kyoto, 616‐8354 Japan;
3. Department of International Agricultural Development, Tokyo University of Agriculture, Sakuragaoka 1‐1‐1, Setagoya, Tokyo 156‐8902, Japan