1. Graduate School of Life Sciences, Tohoku University, Aoba, Sendai 980‐8578, Japan;
2. Present address: Department of Biological Sciences, Macquarie University, Sydney, NSW 2109, Australia
3. Department of International Agriculture Development, Tokyo University of Agriculture, Sakuragaoka 1‐1‐1, Setagaya, Tokyo 156‐8502, Japan;