1. Graduate School of Global Environmental Studies; Kyoto University; Sakyo-ku; Kyoto; 606-8501; Japan
2. Primate Research Institute; Kyoto University; Sakyo-ku; Kyoto; 606-8501; Japan
3. Laboratory of Systematic Zoology; Graduate School of Science; Kyoto University; Sakyo-ku; Kyoto; 606-8501; Japan
4. Faculty of Education; Shiga University; Shiga 520-0862; Japan
5. Laboratory for Molecular Developmental Biology; Graduate School of Science; Kyoto University; Sakyo-ku; Kyoto; 606-8501; Japan