1. Laboratory Center of Life Sciences, College of Life Sciences; Nanjing Agricultural University; Nanjing 210095 China
2. Institute of Botany, Jiangsu Province and the Chinese Academy of Sciences; Jiangsu Province Key Laboratory for Plant Ex-Situ Conservation; Nanjing 210014 China
3. MOA, Laboratory of Plant Nutrition and Fertilization in Lower-Middle Reaches of the Yangtze River; Nanjing Agricultural University; Nanjing 210095 China
4. Centre for Plant Sciences, School of Biology, Faculty of Biological Sciences; University of Leeds; Leeds LS2 9JT UK