1. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐Bioresources South China Agricultural University Guangzhou 510642 China
2. College of Life Sciences South China Agricultural University Guangzhou 510642 China
3. Department of Grassland Science College of Forestry and Landscape Architecture, South China Agricultural University Guangzhou 510642 China
4. Guangdong Engineering Research Center for Grassland Science Guangzhou 510642 China
5. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, South China Agricultural University Guangzhou 510642 China
6. Guangdong Provincial Key Laboratory of Protein Function and Regulation in Agricultural Organisms, College of Life Sciences, South China Agricultural University Guangzhou 510642 China