1. The James Hutton Institute Craigiebuckler Aberdeen AB15 8QH UK
2. The James Hutton Institute Invergowrie Dundee DD2 5DA UK
3. Center for Resources, Environment and Food Security College of Resources and Environmental Sciences China Agricultural University Key Laboratory of Plant and Soil Interactions Ministry of Education Beijing 100193 China
4. Institute of Biological and Environmental Sciences University of Aberdeen Aberdeen AB24 3UU UK
5. Division of Plant Sciences University of Dundee at The James Hutton Institute Invergowrie Dundee DD2 5DA UK
6. School of Plant Biology University of Western Australia Crawley WA 6009 Australia
7. Institute of Evolutionary Biology and Environmental Studies University of Zürich Winterthurerstrasse 190 CH‐8057 Zürich Switzerland
8. SRUC Craibstone Estate Aberdeen AB21 9YA UK