1. State Key Laboratory of Genetic Resources and Evolution; Kunming Institute of Zoology, Chinese Academy of Sciences; Kunming Yunnan 650223 China
2. Department of Geography; Salem State University; Salem MA 01970 U.S.A.
3. State Key Laboratory of Environmental Criteria and Risk Assessment; Chinese Research Academy of Environmental Sciences; Beijing 100012 China
4. China Program; The Nature Conservancy; Kunming Yunnan 650031 China
5. Yunnan Key Laboratory of International Rivers and Transboundary Eco-Security; Yunnan University; Kunming Yunnan China
6. School of Biological Sciences; University of East Anglia, Norwich Research Park; Norwich Norfolk NR47TJ United Kingdom