1. Ocean Biogeochemistry and EcosystemsNational Oceanography Centre Southampton SO14 3ZH U.K.
2. University of Southampton Southampton SO14 3ZH U.K.
3. University of Hawaii Honolulu HI 96822 U.S.A.
4. Centre for Environment Fisheries and Aquaculture Science Lowestoft NR33 0HT U.K.