1. Key Laboratory of Biodiversity and Biogeography; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming 650201 China
2. Department of Botany; Field Museum of Natural History; Chicago IL 60605 USA
3. Plant Germplasm and Genomics Center; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming 650201 China
4. Royal Botanic Garden Edinburgh; Edinburgh EH3 5LR UK