1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming 650201 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Center for Integrative Conservation; Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences; Mengla 666303 China
4. Institute of Systematic Botany; Zollikerstrasse 107 8008 Zurich Switzerland
5. Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming 650201 China