1. Centre for Biodiversity Dynamics Department of Biology Norwegian University of Science and Technology Trondheim Norway
2. Ecology and Genetics Research Unit University of Oulu Oulu Finland
3. Research Unit of Mathematical Sciences University of Oulu Oulu Finland
4. Biocenter Oulu University of Oulu Finland
5. Norwegian Institute for Nature Research Trondheim Norway
6. Department of Ecology and Genetics Uppsala University Uppsala Sweden
7. Infotech Oulu University of Oulu Oulu Finland