1. Institute of Biological and Environmental Sciences; University of Aberdeen; Aberdeen UK
2. State Key Laboratory of Molecular Developmental Biology; Institute of Genetics and Developmental Biology; Chinese Academy of Sciences; Chaoyang Beijing China
3. Key Laboratory of Systems Biology, Innovation Center for Cell Signaling Network; Institute of Biochemistry and Cell Biology; Shanghai Institute of Biological Sciences; Chinese Academy of Sciences; Shanghai China
4. Key Laboratory of Computational Biology; Chinese Academy of Sciences-Max Planck Partner Institute for Computational Biology; Shanghai Institutes for Biological Sciences; Chinese Academy of Sciences; Shanghai China
5. Department of Pathology and Department of Biology; University of Washington; Seattle WA USA