1. School of Biological and Chemical Sciences Queen Mary, University of London London E1 4NS UK
2. Centre for Ocean Life National Institute for Aquatic Resources Technical University of Denmark Kavalergården 6 Charlottenlund 2920 Denmark
3. Department of Biology Juniata College Huntingdon 16652 PA USA
4. Institute of Integrative Biology University of Liverpool Biosciences Building Crown Street Liverpool L69 7ZB UK