1. Division of Immunology; Department of Infectious Diseases and Immunology; Utrecht University; the Netherlands
2. Musculoskeletal Research Group; Institute of Cellular Medicine; Newcastle University; Newcastle upon Tyne UK
3. Arthritis Research UK Rheumatoid Arthritis Pathogenesis Centre of Excellence (RACE); UK UK
4. NIHR-Newcastle Biomedical Research Centre in Ageing and Long-Term Conditions; Newcastle upon Tyne Hospitals NHS Foundation Trust and Newcastle University; Newcastle upon Tyne UK
5. Department of Rheumatology & Clinical Immunology; UMC Utrecht; Utrecht the Netherlands