1. Institute of Hydrobiology and Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms Guangdong Higher Education Institutes Jinan university Guangzhou China
2. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai) Guangzhou China
3. Zhuhai Key Laboratory of Marine Bioresources and Environment Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering School of Marine Sciences Sun Yat‐Sen University Guangzhou China