1. Key Laboratory of Conservation Biology for Endangered Wildlife of the Ministry of Education, and College of Life Sciences Zhejiang University Hangzhou 310058 China
2. Laboratory of Systematic & Evolutionary Botany and Biodiversity Institute of Plant Sciences, and Conservation Center for Gene Resources of Endangered Wildlife Zhejiang University Hangzhou China
3. Department of Organismic Biology Salzburg University A‐5020 Salzburg Austria
4. Department of Biodiversity and Plant Systematics and Botanical Garden and Herbarium Heidelberg (HEID) Center for Organismal Studies (COS) Heidelberg Heidelberg University Heidelberg Germany