1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology, Institute of Biodiversity, School of Ecology and Environmental Science, Yunnan University, Kunming 650500, China
2. Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun 666303, China
3. Yunnan International Joint Laboratory of Southeast Asia Biodiversity Conservation, Menglun 666303, China
4. The Key Laboratory of Rare and Endangered Forest Plants of National Forestry and Grassland Administra-tion & The Key Laboratory for Silviculture and Forest Resources Development of Yunnan Province, Yun-nan Academy of Forestry and Grassland, Kunming 650201, China