1. Plant Conservation and Population Biology, Biology Dept, KU Leuven Leuven Belgium
2. Inst. of Ecology and Evolutionary Biology, National Taiwan Univ., Da'an District Taipei Taiwan
3. Research Faculty of Agriculture, Hokkaido Univ. Kita‐ku Sapporo Japan
4. Division of Forest, Nature and Landscape, KU Leuven Leuven Belgium