1. Jiangsu Key Laboratory of Crop Genetics and Physiology/Key Laboratory of Plant Functional Genomics of the Ministry of Education/Jiangsu Key Laboratory of Crop Genomics and Molecular Breeding Agricultural College of Yangzhou University Yangzhou 225009 China
2. Jiangsu Co‐Innovation Center for Modern Production Technology of Grain Crops Yangzhou University Yangzhou 225009 China
3. Key Laboratory of Molecular Epigenetics of the Ministry of Education (MOE) Northeast Normal University Changchun 130024 China
4. Department of Plant Biology Department of Horticulture Michigan State University East Lansing Michigan 48824 USA
5. Joint International Research Laboratory of Agriculture and Agri‐Product Safety, the Ministry of Education of China Yangzhou University Yangzhou 225009 China