1. College of Agriculture, Nanjing Agricultural University, Nanjing, Jiangsu 210095, China
2. College of Life Science, Zhengzhou University, Zhengzhou, Henan 450001, China
3. The Shennong Laboratory/Nation Industrial Innovation Centre for Bio-Breeding/Key Laboratory of Oil Crops in Huang-Huai-Hai Plains, Ministry of Agriculture/Henan Provincial Key Laboratory for Oil Crops Improvement/Institute of Crop Molecular Breeding, Henan Academy of Agricultural Sciences, Zhengzhou, Henan 450002, China