1. Division of Biology; Kansas State University; 116 Ackert Hall Manhattan KS 66506-4901 U.S.A.
2. Department of Forest and Conservation Sciences; Centre for Applied Conservation Biology, University of British Columbia; 3004-2424 Main Mall Vancouver British Columbia V6T 1Z4 Canada
3. Environment Canada; 5421 Robertson Road Delta British Columbia V4K 3N2 Canada