1. Qianyanzhou Ecological Research Station, Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research Chinese Academy of Sciences Beijing China
2. Zhongke‐Ji'an Institute for Eco‐Environmental Sciences Ji'an China
3. USDA Forest Service Pacific Northwest Research Station Corvallis Oregon
4. Key Laboratory of Environment Change and Resources Use in Beibu Gulf, Ministry of Education Guangxi Teachers Education University Nanning China
5. Department of Botany University of Wisconsin Madison Wisconsin
6. Department of Biology Western Oregon University Monmouth Oregon