1. School of Biological Sciences and Centre for Ecology, Evolution, and Conservation Biology (CEEC), University of East Anglia, Norwich, Norfolk NR4 7TJ, UK;
2. Department of Biology, Imperial College at Silwood Park, Ascot, Berks SL5 7PY, UK;
3. Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA;
4. Herbario Vargas, Universidad Nacional de San Antonio Abad de Cusco, Cusco, Perú
5. Sachavacayoc, Tambopata, Madre de Dios, Perú