1. National Education Ministry, Key Laboratory of Plant Genetic Improvement and Comprehensive Utilization, Fujian Agriculture and Forestry University; Fuzhou Fujian 350002 China
2. College of Crop Science; Fujian Agriculture and Forestry University; Fuzhou Fujian 350002 China
3. College of Life Science; Fujian Agriculture and Forestry University; Fuzhou Fujian 350002 China