1. State Key Laboratory of Cotton Biology, Laboratory of Plant Germplasm and Genetic Engineering, School of Life Sciences Henan University Kaifeng 475000 China
2. Xianrendong Nature Reserve Zhuanghe 116407 China
3. Department of Environment and Forest Resources Chungnam National University Daejeon 34134 South Korea
4. Key Laboratory of Conservation Biology for Endangered Wildlife of the Ministry of Education, and Laboratory of Systematic & Evolutionary Botany and Biodiversity, College of Life Sciences Zhejiang University Hangzhou 310058 China